Soon after the Bills traded Marcell Dareus to the Jacksonville Jaguars last season, the run defense suffered.
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Hours later, a five-year deal was struck between the Buffalo Bills and Lotulelei, a source with direct knowledge of the deal told the Observer. Of those yards, 1,105 were inside the tackles, and opponents' 4.8 yards per rush inside the tackles was tied for worst in the NFL.
The very first draft pick of former Panthers general manager Dave Gettleman, Lotulelei was selected 14th overall in 2013 out of the University of Utah. Williams, 34, has been contemplating his playing future this offseason.
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Lotulelei will be reunited with former Panthers defensive coordinator Sean McDermott, who led Buffalo to its first playoff appearance since the 1999 season in 2017. Past year he had 1.5 sacks and 25 total tackles for Carolina.
O'Leary caught a career-high 22 passes for 322 yards and two touchdowns last season in 15 games, including five starts.
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Lotulelei has recorded 141 tackles and 11.5 sacks in his career.